Crowd groups seemingly ignored when used for controlling remote repo permissions

When Crowd user groups are used to control permissions for a remote repo, Artifactory seems to ignore those settings, even though adding permissions for individual Crowd users works fine.

We are running an old version of Artifactory (working on upgrading, but a lot has changed and it's complicated now), but a search across all other Jira tickets here shows no other match, so there is no reason to believe it's been fixed since v4.10.